ERRO SPRING/STRUTS - HTTP Status 404 - Servlet action is not available

Estou com o seguinte erro (que aparece no browser):
HTTP Status 404 - Servlet action is not available

Quando rodo somente com struts fica ok, mas quando adiciono spring (action-servlet.xml, applicationContext.xml), referenciando no struts-config.xml dá este erro.

Alguém sabe o que é? Segue abaixo meu código.

web.xml

<?xml version="1.0" encoding="ISO-8859-1"?>
<!DOCTYPE web-app PUBLIC "-//Sun Microsystems, Inc.//DTD Web Application 2.3//EN"
                         "http://java.sun.com/dtd/web-app_2_3.dtd">
<web-app>
 <display-name>appdaca-taciano_romulo</display-name>
 <!-- listener>
  <listener-class>ojato.util.ContextListener</listener-class>
 </listener-->
 <!-- Action Servlet Configuration -->
 <servlet>
  <servlet-name>action</servlet-name>
  <servlet-class>org.apache.struts.action.ActionServlet</servlet-class>
  <init-param>
   <param-name>config</param-name>
   <param-value>/WEB-INF/struts-config.xml</param-value>
  </init-param>
  <load-on-startup>1</load-on-startup>
 </servlet>
 <!-- Spring -->
 <!-- servlet>
  <servlet-name>SpringContextServlet</servlet-name>
  <servlet-class>org.springframework.web.context.ContextLoaderServlet</servlet-class>
  <load-on-startup>1</load-on-startup>
 </servlet-->
 <!-- Action Servlet Mapping -->
 <servlet-mapping>
  <servlet-name>action</servlet-name>
  <url-pattern>*.do</url-pattern>
 </servlet-mapping>
 <!-- The Usual Welcome File List -->
 <welcome-file-list>
  <welcome-file>index.jsp</welcome-file>
 </welcome-file-list>
 <taglib>
  <taglib-uri>/WEB-INF/struts-bean</taglib-uri>
  <taglib-location>/WEB-INF/struts-bean.tld</taglib-location>
 </taglib>
 <taglib>
  <taglib-uri>/WEB-INF/struts-logic</taglib-uri>
  <taglib-location>/WEB-INF/struts-logic.tld</taglib-location>
 </taglib>
 <taglib>
  <taglib-uri>/WEB-INF/struts-html</taglib-uri>
  <taglib-location>/WEB-INF/struts-html.tld</taglib-location>
 </taglib>
 <login-config>
  <auth-method>BASIC</auth-method>
 </login-config>
</web-app>

struts-config.xml

<?xml version="1.0" encoding="UTF-8"?>
<!DOCTYPE struts-config PUBLIC "-//Apache Software Foundation//DTD Struts Configuration 1.1//EN"
                               "http://jakarta.apache.org/struts/dtds/struts-config_1_1.dtd">
<struts-config>
 <data-sources/>
 <form-beans>
  <!-- form-bean name="GetNameForm" type="demo.GetNameForm" / -->
  <form-bean name="clienteForm" type="ojato.cliente.controller.ClienteForm"/>
 </form-beans>
 <global-exceptions/>
 <global-forwards>
  <!-- forward name="getName" path="/pages/inputname.jsp"/ -->
  <forward name="paginaInicial" path="/pages/index.html"/>
 </global-forwards>
 <action-mappings>
  <!-- action name="GetNameForm" path="/greeting" scope="request" type="demo.GreetingAction">
   <forward name="sayhello" path="/pages/greeting.jsp"/>
  </action-->
  <action name="clienteForm" parameter="metodo" path="/cliente"
   scope="request"
   type="org.springframework.web.struts.DelegatingActionProxy" validate="false">
   <forward name="editarCliente" path="/pages/editarCliente.jsp"/>
  </action>
 </action-mappings>
 <controller/>
 <plug-in className="org.springframework.web.struts.ContextLoaderPlugIn">
  <set-property property="contextConfigLocation" value="/WEB-INF/applicationContext.xml, /WEB-INF/action-servlet.xml"/>
 </plug-in>
 <plug-in className="org.apache.struts.validator.ValidatorPlugIn">
  <set-property property="pathnames" value="/WEB-INF/validator-rules.xml, /WEB-INF/validation.xml"/>
 </plug-in>
</struts-config>

action-servlet.xml

<?xml version="1.0" encoding="UTF-8"?>
<!DOCTYPE beans PUBLIC "-//SPRING//DTD BEAN//EN"
                       "http://www.springframework.org/dtd/spring-beans.dtd">
<beans>
 <bean class="ojato.cliente.controller.ClienteAction" name="/cliente" singleton="false">
  <property name="manipuladorCliente">
   <ref bean="manipuladorCliente"/>
  </property>
 </bean>
</beans>

applicationContext.xml

<?xml version="1.0" encoding="UTF-8"?>
<!DOCTYPE beans PUBLIC "-//SPRING//DTD BEAN//EN"
"http://www.springframework.org/dtd/spring-beans.dtd">

<beans>
	<bean id="daoCliente"
		class="ojato.cliente.database.dao.HibernateClienteDao">
		<property name="sessionFactory">
			<ref local="sessionFactory" />
		</property>
	</bean>

	<bean id="manipuladorCliente"
		class="ojato.cliente.logic.ManipuladorCliente">
		<property name="daoCliente">
			<ref bean="daoCliente" />
		</property>
	</bean>

	<bean id="sessionFactory"
		class="org.springframework.orm.hibernate3.LocalSessionFactoryBean">
		<property name="dataSource">
			<ref local="dataSource" />
		</property>
		<property name="mappingResources">
			<value>ojato/cliente/database/Cliente.hbm.xml,</value>
		</property>
		<property name="hibernateProperties">
			<props>
				<prop key="hibernate.dialect">
					org.hibernate.dialect.MySQLDialect
				</prop>
				<prop key="show_sql">true</prop>
				<prop key="hibernate.generate_statistics">true</prop>
				<prop key="hibernate.use_sql_comments">true</prop>
				<prop key="hibernate.cache.provider_class">
					org.hibernate.cache.EhCacheProvider
				</prop>
				<prop key="hibernate.cglib.use_reflection_optimizer">
					false
				</prop>
			</props>
		</property>

	</bean>


	<bean id="dataSource"
		class="org.apache.commons.dbcp.BasicDataSource"
		destroy-method="close">
		<property name="url">
			<value>jdbc:mysql://localhost:3306/appdaca</value>
		</property>
		<property name="username">
			<value>appdaca</value>
		</property>
		<property name="password">
			<value>appdaca</value>
		</property>
		<property name="driverClassName">
			<value>com.mysql.jdbc.Driver</value>
		</property>
		<property name="initialSize">
			<value>1</value>
		</property>
		<property name="maxActive">
			<value>2</value>
		</property>
		<property name="maxIdle">
			<value>2</value>
		</property>
		<property name="poolPreparedStatements">
			<value>true</value>
		</property>
	</bean>

</beans>

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